Tropaeolum – Nasturtium

Balcony grown in a small hanging basket from seed.

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Tropaeolum /trɵˈpiːələm/, commonly known as nasturtium (/næˈstɜrʃ(i)əm/; literally “nose-twister” or “nose-tweaker”), is a genus of roughly 80 species of annual and perennial herbaceous flowering plants. It was named by Carl Linnaeus and is the only genus in the family Tropaeolaceae. The nasturtiums received their common name because they produce an oil that is similar to that of watercress (Nasturtium officinale).

The genus Tropaeolum, native to South and Central America, includes several very popular garden plants, the most commonly grown being T. majus, T. peregrinum and T. speciosum. One of the hardiest species is T. polyphyllum from Chile, the perennial roots of which can survive the winter underground at altitudes of 3,300 metres (10,000 ft).

Plants in this genus have showy, often intensely bright flowers, and rounded, peltate (shield-shaped) leaves with the petiole in the centre. The flowers are bisexual and zygomorphic, with five petals, a superior three-carpelled ovary, and a funnel-shaped nectar spur at the back, formed by modification of one of the five sepals.

Kingdom:	Plantae
(unranked):	Angiosperms
(unranked):	Eudicots
(unranked):	Rosids
Order:	Brassicales
Family:	Tropaeolaceae
Genus:	Tropaeolum L.

History

The first Tropaeolum species was imported into Spain by the Spanish botanist Nicolás Monardes. He published an account in 1569 entitled Joyful News out of the Newe Founde Worlde in which he described, among other things, the plants and animals discovered in South America. The English herbalist John Gerard reports having received seeds of the plant from Europe in his 1597 book Herball, or Generall Historie of Plantes. Tropaeolum majus was named by the Swedish botanist Carl Linnaeus, who chose the genus name because the plant reminded him of an ancient custom. After victory in battle, the Romans used to set up a trophy pole called a tropaeum (from the Greek tropaion, source of English “trophy“). On this the armour and weapons of the vanquished foe were hung. Linnaeus was reminded of this by the plant as the round leaves resembled shields and the flowers, blood-stained helmets.

Nasturtiums were once known commonly as “Indian cresses” because they were introduced from the Americas, known popularly then as the Indies, and used like cress as salad ingredients. John Gerard called the plant “Indian Cresses” in his herbal. He wrote: “unto the backe part (of the flower) doth hange a taile or spurre, such as hath the Larkes heele, called in Latine Consolida regalis. He was comparing the flowers of Indian cress to those of forking larkspur (Consolida regalis) of the buttercup family. J R R Tolkien commented that an alternative anglicization of the name was “nasturtian” rather than “nasturtium”.

Description

Tropaeolum is a genus of dicotyledonous annual or perennial plants, often with somewhat succulent stems and sometimes tuberous roots. The alternate leaves are hairless, peltate and entire or palmately lobed. The petioles or leaf stalks are long and in many species are able to twine round other stems to provide support. The flowers are bisexual and showy, set singly on long stalks in the axils of the leaves. They have five sepals, the uppermost of which is elongated into a nectar spur. The five petals are clawed, with the lower three unlike the upper two. The eight stamens are in two whorls of unequal length and the superior ovary has three segments and three stigmas on a single style. The fruit is naked and nut-like, with three single seed segments.

Tropaeolum tuberosum

The most common flower in cultivation is a hybrid of T. majus, T. minus and T. peltophorum, and is commonly known as the nasturtium (and occasionally anglicized as nasturtian). It is mostly grown from seed as a half-hardy annual and both single and double varieties are available. It comes in a range of forms and colours including cream, yellow, orange and red, solid in colour or striped and often with a dark blotch at the base of the petals. It is vigorous and easily grown and does well in sun. It thrives in poor soil and dry conditions, whereas in rich soil it tends to produce much leafy growth and few flowers. Some varieties adopt a bush form while others scramble over and through other plants and are useful for planting in awkward spots or for covering fences and trellises.

The following cultivars have gained the Royal Horticultural Society‘s Award of Garden Merit:

• ‘Alaska Series’
• ‘Hermine Grashoff’
• ‘Whirlybird Series’

The blue nasturtium (Tropaeolum azureum) is a tender species from Chile which has violet-blue flowers with white eyes that can be as much as 4 cm (1.6 in) across

Tropaeolum brachyceras has 2.5 cm (1 in) yellow flowers with purplish markings on wiry, climbing stems. It is a half hardy perennial from Chile and may remain dormant for several years before being sparked into growth by some unknown trigger factor.

Tropaeolum hookerianum is a tuberous-rooted species from Chile. There are two subspecies, T. h. austropurpureum which has violet-purple flowers and T. h. pilosum with yellow flowers.

The Canary creeper (Tropaeolum peregrinum) is a trailing and climbing half-hardy annual species with wiry stalks and palmately lobed leaves. The pale yellow, fringed flowers are borne on long stalks. It originated from Peru but may first have been cultivated in the Canary Islands before being introduced into Western Europe.

Wreath nasturtium (Tropaeolum polyphyllum) is a prostrate plant originating from Argentina and Chile. It has silvery, deeply lobed leaves and a profusion of small, bright yellow flowers on long trailing stalks. After flowering, the plant dies back. It is a perennial with underground rhizomes which send up new shoots at intervals. In a suitable sunny location with well drained soil, it will survive for several years. It is a very hardy species; the tubers can grow at depths of 60 cm (24 in) enabling the plant to survive at altitudes of as much as 3,300 metres (10,000 ft) in the Andes.

The flame flower (Tropaeolum speciosum) is well adapted to cool, moist climates and notoriously does well in Scotland. It sends up shoots which thread their way through hedges and shrubs and which, when they emerge into the light, bear brilliant red flowers among small, five or six-lobed leaves. It is difficult to establish but is an attractive garden plant when it thrives. This plant has gained the Royal Horticultural Society’s Award of Garden Merit.

Three-coloured Indian cress (Tropaeolum tricolor) is another tuberous, climbing species grown for its attractive red, purple and yellow tubular flowers. It comes from Chile and Bolivia and is a reliable winter-growing species.

Mashua (Tropaeolum tuberosum) is a perennial climbing plant from the Andes grown for its tuberous roots. It has been cultivated since ancient times and depictions of it are found at archaeological sites pre-dating the Incas. It has leaves with five to seven lobes and small, long-spurred, red and yellow flowers. The tubers have an unpleasant smell when raw which disappears on cooking. It is frost-hardy and produces crops of 30 tons per hectare at a height of 3,000 metres (9,800 ft) above sea level. The cultivar T. tuberosum lineamaculatum ‘Ken Aslet’ has gained the Royal Horticultural Society’s Award of Garden Merit.

Species that originated from the coastal areas and from the lower foothills make most of their growth in winter whereas the true alpine species are summer growers. Tuberous Tropaeolum species are well known for occasionally remaining dormant for one or more years. The species with underground rhizomes and tubers can be propagated from these while other species are best raised from seed. Fresh seed is favoured by many growers but dried seed is also often successful. Seed from the winter growing species should be sown in the autumn while the summer growing species are best sown in the spring in well-drained compost and covered with 1 to 2 cm (0.4 to 0.8 in) of grit or sand. The containers should be kept at below 4 °C (39 °F) until the seedlings appear in about a month, as too high a temperature inhibits germination.

Uses

Culinary

All parts of T. majus are edible. The flower has most often been consumed, making for an especially ornamental salad ingredient; it has a slightly peppery taste reminiscent of watercress, and is also used in stir fry. The flowers contain about 130 mg vitamin C per 100 grams (3.5 oz)^] about the same amount as is contained in parsley.

Moreover, they contain up to 45 mg of lutein per 100 gr, which is the highest amount found in any edible plant. The unripe seed pods can be harvested and dropped into spiced vinegar to produce a condiment and garnish, sometimes used in place of capers.

Mashua (T. tuberosum) produces an edible underground tuber that is a major food source in parts of the Andes.

Herbal medicine

Nasturtiums have been used in herbal medicine for their antiseptic and expectorant qualities. They are said to be good for chest colds and to promote the formation of new blood cells. T. majus has been used in herbal medicine for respiratory and urinary tract infections.

Taxonomy

Tropaeolum was previously placed in the family Tropaeolaceae along with two other genera, Magallan and Trophaeastrum. The monotypic genus Magallan was characterised by having winged fruit, and the two species of Trophaeastrum lacked spurs. The genus Tropaeolum was diagnosed only by the absence of the characteristics of the other two genera. A molecular study undertaken in 2000 found Tropaeolum to be paraphyletic when the other two genera are segregated, so Magallan and Trophaeastrum were reduced to synonyms of Tropaeolum. Tropaeolaceae was thus rendered monogeneric, a family of only one genus.

Species

“The Plant List”, a collaboration between the Missouri Botanical Garden and the Royal Botanic Gardens, Kew includes the following accepted names of Tropaeolum species names. Some that are under review, are here marked “U”.

Tropaeolum adpressum Tropaeolum albiflorum – U Tropaeolum argentinum Tropaeolum asplundii Tropaeolum atrocapillare Tropaeolum atrosanguineum – U Tropaeolum azureum Tropaeolum beuthii Tropaeolum bicolor Tropaeolum bogotense – U Tropaeolum boliviense – U Tropaeolum brachyceras Tropaeolum brasiliense Tropaeolum brideanum Tropaeolum buchenavianum – U Tropaeolum calcaratum Tropaeolum calvum Tropaeolum capillare Tropaeolum carchense Tropaeolum ciliatum Tropaeolum cirrhipes Tropaeolum cochabambae Tropaeolum concavum Tropaeolum concinneum – U Tropaeolum canariense Tropaeolum crenatiflorum Tropaeolum cubio – U Tropaeolum curvirostre – U Tropaeolum cuspidatum Tropaeolum dawei – U Tropaeolum deckerianum Tropaeolum denticualtum – U

Tropaeolum dicolorum – U Tropaeolum dipetalum Tropaeolum elzae – U Tropaeolum emarginatum Tropaeolum ferreyrae Tropaeolum fintelmannii Tropaeolum flavipilum – U Tropaeolum garciae – U Tropaeolum harlingii Tropaeolum hayneanum Tropaeolum hirsutum – U Tropaeolum hirtifolium – U Tropaeolum hjertingii Tropaeolum hookerianum – U Tropaeolum hughesae – U Tropaeolum huigrense Tropaeolum incisum Tropaeolum integrifolium – U Tropaeolum jarrattii – U Tropaeolum jilesii Tropaeolum karstenii – U Tropaeolum kerneisinum Tropaeolum killipii – U Tropaeolum kingii Tropaeolum klotzschii – U Tropaeolum kuntzeanum Tropaeolum lasseri Tropaeolum lehmannii – U Tropaeolum leichtlinii – U Tropaeolum leonis Tropaeolum leptophyllum Tropaeolum lindenii – U Tropaeolum longiflorum

Tropaeolum longifolium – U Tropaeolum looseri Tropaeolum macrophyllum – U Tropaeolum maculifolium – U Tropaeolum magnificum Tropaeolum majus Tropaeolum marginatum Tropaeolum menispermifolium Tropaeolum mexiae – U Tropaeolum meyeri Tropaeolum minus Tropaeolum moritzianum Tropaeolum morreanum – U Tropaeolum myriophyllum Tropaeolum naudinii – U Tropaeolum nuptae-jucundae Tropaeolum orinocense Tropaeolum orthoceras – U Tropaeolum oxalidanthum – U Tropaeolum papillosum Tropaeolum parvifolium – U Tropaeolum pellucidum – U Tropaeolum peltophorum Tropaeolum pendulum Tropaeolum pentagonum Tropaeolum pentaphyllum Tropaeolum peregrinum Tropaeolum polyphyllum Tropaeolum popelari – U Tropaeolum porifolium Tropaeolum pubescens Tropaeolum purpureum

Tropaeolum reichianum – U Tropaeolum reineckeanum – U Tropaeolum repandum – U Tropaeolum rhizophorum – U Tropaeolum rhomboideum Tropaeolum sanctae-catharinae Tropaeolum scheuerianum – U Tropaeolum schillingii – U Tropaeolum schlimii – U Tropaeolum seemannii Tropaeolum sessilifolium Tropaeolum smithii Tropaeolum sparrei – U Tropaeolum speciosum Tropaeolum steyermarkianum Tropaeolum stipulatum Tropaeolum tenellum – U Tropaeolum tenuirostre – U Tropaeolum traceyae – U Tropaeolum trialatum Tropaeolum trialatum Tropaeolum tricolor Tropaeolum tricolori-brachyceras – U Tropaeolum trilobum Tropaeolum trilobum Tropaeolum tuberosum Tropaeolum umbellatum Tropaeolum unilobatum – U Tropaeolum wagnerianum Tropaeolum warmingianum Tropaeolum warscewiczii – U Tropaeolum willinkii

Taraxacum officinale – Dandelion

Taraxacum officinale, the common dandelion (often simply called “dandelion“), is a flowering herbaceous perennial plant of the family Asteraceae (Compositae). It can be found growing in temperate regions of the world, in lawns, on roadsides, on disturbed banks and shores of water ways, and other areas with moist soils. T. officinale is considered a weed, especially in lawns and along roadsides, but it is sometimes used as a medical herb and in food preparation. Common dandelion is well known for its yellow flower heads that turn into round balls of silver tufted fruits that disperse in the wind called “blowballs”^[3] or “clocks” (in both British and American English).

Scientific classification
Kingdom:	Plantae
(unranked):	Angiosperms
(unranked):	Eudicots
(unranked):	Asterids
Order:	Asterales
Family:	Asteraceae
Tribe:	Cichorieae
Genus:	Taraxacum
Species:	T. officinale

Description

Taraxacum officinale grows from generally unbranched taproots and produces one to more than ten stems that are typically 5 to 40 cm tall, but sometimes up to 70 cm tall. The stems can be tinted purplish, they are upright or lax, and produce flower heads that are held as tall or taller than the foliage. The foliage may be upright-growing or horizontally spreading; the leaves have petioles that are either unwinged or narrowly winged. The stems can be glabrous or sparsely covered with short hairs. Plants have milky latex and the leaves are all basal; each flowering stem lacks bracts and has one single flower head. The yellow flower heads lack receptacle bracts and all the flowers, which are called florets, are ligulate and bisexual. The fruits are mostly produced by apomixis.

The leaves are 5 to 45 cm long and 1 to 10 cm wide, and are oblanceolate, oblong, or obovate in shape, with the bases gradually narrowing to the petiole. The leaf margins are typically shallowly lobed to deeply lobed and often lacerate or toothed with sharp or dull teeth.

The calyculi (the cuplike bracts that hold the florets) are composed of 12 to 18 segments: each segment is reflexed and sometimes glaucous. The lanceolate shaped bractlets are in two series, with the apices acuminate in shape. The 14- to 25-mm wide involucres are green to dark green or brownish-green, with the tips dark gray or purplish. The florets number 40 to over 100 per head, having corollas that are yellow or orange-yellow in color.

The fruits, called cypselae, range in color from olive-green or olive-brown to straw-colored to grayish, they are oblanceoloid in shape and 2 to 3 mm long with slender beaks. The fruits have 4 to 12 ribs that have sharp edges. The silky pappi, which form the parachutes, are white to silver-white in color and around 6 mm wide. Plants typically have 24 or 40 pairs of chromosomes but some plants have 16 or 32 chromosomes.

Taxonomy

The taxonomy of the genus Taraxacum is complicated by apomictic and polyploid lineages, and the taxonomy and nomenclatural situation of Taraxacum officinale is not yet fully resolved, The taxonomy of this species has in the past been complicated by the recognition of numerous species, subspecies and microspecies. E.g. Rothmaler’s flora of Germany recognizes roughly 70 microspecies.The plants introduced to North America are triploids that reproduce by obligate gametophytic apomixis Some authorities recognize three subspecies of Taraxacum officinale including:

• Taraxacum officinale ssp. ceratophorum (Ledeb.) Schinz ex Thellung which is commonly called common dandelion, fleshy dandelion, horned dandelion or rough dandelion. It is native to Canada and the western US. Some sources list it as a species, Taraxacum ceratophorum.
• Taraxacum officinale ssp. officinale, which is commonly called common dandelion or wandering dandelion.
• Taraxacum officinale ssp. vulgare (Lam.) Schinz & R. Keller, which is commonly called common dandelion.

Two of them have been introduced and established in Alaska and the third (ssp. ceratophorum ) is native there.

Taraxacum officinale has many English common names (of which some are no longer in use), including blowball, lion’s-tooth, cankerwort, milk-witch, yellow-gowan, Irish daisy, monks-head, priest’s-crown and puff-ball; other common names include, faceclock, pee-a-bed, wet-a-bed,  swine’s snout,  white endive, and wild endive.

Carl Linnaeus named the species Leontodon Taraxacum in 1753. The genus name Taraxacum, might be from the Arabic word “Tharakhchakon“, or from the Greek word “Tarraxos“. The common name “dandelion,” comes from the French phrase “dent de lion” which means “lion’s tooth”, in reference to the jagged shaped foliage.

Ecology

Taraxacum officinale is native to Eurasia, and now is naturalized throughout North America, southern Africa, South America, New Zealand, Australia, and India. It occurs in all 50 states of the USA and most Canadian provinces. It is considered a noxious weed in some jurisdictions, and is considered to be a nuisance in residential and recreational lawns in North America. It is also an important weed in agriculture and causes significant economic damage because of its infestation in many crops worldwide.

The dandelion is a common colonizer of disturbed habitats, both from wind blown seeds and seed germination from the seed bank. The seeds remain viable in the seed bank for many years, with one study showing germination after nine years. This species is a somewhat prolific seed producer, with 54 to 172 seeds produced per head, and a single plant can produce more than 5,000 seeds a year. It is estimated that more than 97,000,000 seeds/hectare could be produced yearly by a dense stand of dandelions^.  When released, the seeds can be spread by the wind up to several hundred meters from their source. The seeds are also a common contaminant in crop and forage seeds. The plants are adaptable to most soils and the seeds are not dependent on cold temperatures before they will germinate but they need to be within the top 2.5 centimeters of soil.

While not in bloom, this species is sometimes confused with others, such as Chondrilla juncea, that have similar basal rosettes of foliage.  Another plant, sometimes referred to as Fall Dandelion, is very similar to dandelion, but produces “yellow fields” later.

Uses

While the dandelion is considered a weed by many gardeners and lawn owners, the plant has several culinary and medicinal uses. The specific name officinalis refers to its value as a medicinal herb, and is derived from the word opificina, later officina, meaning a workshop or pharmacy. The flowers are used to make dandelion wine, the greens are used in salads, the roots have been used to make a coffee substitute (when baked and ground into powder) and the plant was used by Native Americans as a food and medicine.

Culinary

Dandelions are wildcrafted or grown on a small scale as a leaf vegetable. The leaves (called dandelion greens) can be eaten cooked or raw in various forms, such as in soup or salad. They are probably closest in character to mustard greens. Usually the young leaves and unopened buds are eaten raw in salads, while older leaves are cooked. Raw leaves have a slightly bitter taste. Dandelion salad is often accompanied with hard boiled eggs. The leaves are high in vitamin A, vitamin C and iron, carrying more iron and calcium than spinach.

Dandelion flowers can be used to make dandelion wine, for which there are many recipes. Most of these are more accurately described as “dandelion-flavored wine,” as some other sort of fermented juice or extract serves as the main ingredient. It has also been used in a saison ale called Pissenlit (literally “wet the bed” in French) made by Brasserie Fantôme in Belgium. Dandelion and burdock is a soft drink that has long been popular in the United Kingdom.

Another recipe using the plant is dandelion flower jam. In Silesia and also other parts of Poland and world, dandelion flowers are used to make a honey substitute syrup with added lemon (so-called May-honey). This “honey” is believed to have a medicinal value, in particular against liver problems. Ground roasted dandelion root can be used as a non-caffeinated coffee substitute.

Herbal medicine

Dandelion root medicinally is sold primarily  as a diuretic.  A hepatoprotective effect in mice of chemicals extracted from dandelion root has been reported.Dandelion is used in herbal medicine as a mild laxative, for increasing appetite, and for improving digestion. The milky latex has been used as a mosquito repellent and as a folk remedy to treat warts.

Other

Yellow or green dye colours can be obtained from the flowers but little colour can be obtained from the roots of the plant.

T. officinale is food for the caterpillars of several Lepidoptera (butterflies and moths), such as the tortrix moth Celypha rufana. See also List of Lepidoptera that feed on dandelions.

Toxicity

Dandelion (Taraxacum officinale) has been linked to outbreaks of stringhalt in horses.

Ranunculus – Buttercup etc.

Ranunculus /ræˈnʌŋkjʊləs/is a large genus of about 600 species of plants in the Ranunculaceae. Members of the genus include the buttercups, spearworts, water crowfoots and the lesser celandine.

Scientific classification
Kingdom: Plantae
(unranked): Angiosperms
(unranked): Eudicots
Order: Ranunculales
Family: Ranunculaceae
Genus: Ranunculus

They are mostly herbaceous perennials with bright yellow or white flowers (if white, still with a yellow centre); some are annuals or biennials. A few species have orange or red flowers. There are usually five petals, but sometimes six, numerous, or none, as in R. auricomus. The petals are often highly lustrous, especially in yellow species. Buttercups usually flower in the spring, but flowers may be found throughout the summer, especially where the plants are growing as opportunistic colonizers, as in the case of garden weeds.

The Water crowfoots (Ranunculus subgenus Batrachium), which grow in still or running water, are sometimes treated in a separate genus Batrachium (from Greek batrachos βάτραχος: frog, etymology comparable to ranunculus itself). They have two different leaf types, thread-like leaves underwater and broader floating leaves. In some species, such as R. aquatilis, a third, intermediate leaf type occurs.

Ranunculus species are used as food plants by the larvae of some Lepidoptera species including Hebrew Character and Small Angle Shades. Some species are popular ornamental flowers in horticulture, with many cultivars selected for large and brightly coloured flowers

Naming

The name Ranunculus is Late Latin for “little frog,” from rana “frog” and a diminutive ending. This probably refers to many species being found near water, like frogs.

The name buttercup may derive from a false belief that the plants give butter its characteristic yellow hue (in fact it is poisonous to cows and other livestock). A popular children’s game involves holding a buttercup up to the chin; a yellow reflection is supposed to indicate fondness for butter.

In the interior of the Pacific Northwest of the United States the buttercup is called “Coyote’s eyes” — ʔiceyéeyenm sílu in Nez Perce and spilyaynmí áčaš in Sahaptin. In the legend Coyote was tossing his eyes up in the air and catching them again when Eagle snatched them. Unable to see, Coyote made eyes from the buttercup.^{[citation needed]}

Splitting of the genus

Molecular investigation of the genus has revealed that Ranunculus is not monophyletic with respect to a number of other recognized genera in the family – e.g. Ceratocephala, Halerpestes, Hamadryas, Laccopetalum, Myosurus, Oxygraphis, Paroxygraphis and Trautvetteria. A proposal to split Ranunculus into several genera have thus been published in a new classification for the tribe Ranunculeae. The split (and often re-recognized) genera include Arcteranthis Greene, Beckwithia Jeps., Callianthemoides Tamura, Coptidium (Prantl) Beurl. ex Rydb., Cyrtorhyncha Nutt. ex Torr. & A.Gray, Ficaria Guett., Krapfia DC., Kumlienia E.Greene and Peltocalathos Tamura.

Toxicity

All Ranunculus species are poisonous when eaten fresh by cattle, horses, and other livestock, but their acrid taste and the blistering of the mouth caused by their poison means they are usually left uneaten. Poisoning can occur where buttercups are abundant in overgrazed fields where little other edible plant growth is left, and the animals eat them out of desperation. Symptoms include bloody diarrhea, excessive salivation, colic, and severe blistering of the mucous membranes and gastrointestinal tract. When Ranunculus plants are handled, naturally occurring ranunculin is broken down to form protoanemonin, which is known to cause contact dermatitis in humans and care should therefore be exercised in extensive handling of the plants.    The toxins are degraded by drying, so hay containing dried buttercups is safe.

Selected species list

• Ranunculus abortivus – littleleaf buttercup
• Ranunculus aconitifolius – aconite-leaf buttercup
• Ranunculus acraeus – a newly described species from Otago, New Zealand
• Ranunculus acris – meadow buttercup
• Ranunculus alismifolius – plantainleaf buttercup
• Ranunculus andersonii – Anderson’s buttercup
• Ranunculus aquatilis – common water crowfoot
• Ranunculus arvensis – corn buttercup
• Ranunculus asiaticus – Persian buttercup
• Ranunculus auricomus – Goldilocks buttercup (type species)
• Ranunculus biternatus – Antarctic buttercup
• Ranunculus bonariensis – Carter’s buttercup
• Ranunculus bulbosus – bulbous buttercup
• Ranunculus calandrinioides – high alpine buttercup
• Ranunculus californicus – California buttercup
• Ranunculus canus – Sacramento Valley buttercup
• Ranunculus cassubicus – Kashubian buttercup
• Ranunculus crassipes – subantarctic buttercup
• Ranunculus cymbalaria – marsh buttercup
• Ranunculus eschscholtzii – alpine buttercup
• Ranunculus ficaria – lesser celandine
• Ranunculus flabellaris – yellow water buttercup
• Ranunculus flammula – lesser spearwort
• Ranunculus fluitans – river water crowfoot
• Ranunculus glaberrimus – sagebrush buttercup
• Ranunculus glacialis – glacier buttercup
• Ranunculus gormanii – Gorman’s buttercup
• Ranunculus hebecarpus – delicate buttercup
• Ranunculus hispidus – bristly buttercup
• Ranunculus hydrocharoides – frogbit buttercup
• Ranunculus jovis – Utah buttercup
• Ranunculus kadzusensis – makino, maehwamarum
• Ranunculus lapponicus – Lapland buttercup
• Ranunculus lingua – greater spearwort
• Ranunculus lobbii – Lobb’s buttercup
• Ranunculus longirostris – water buttercup
• Ranunculus lyallii – Mount Cook lily, reputedly the largest buttercup
• Ranunculus macounii – Macoun’s buttercup
• Ranunculus micranthus – small-flowered crowfoot
• Ranunculus moseleyi – Moseley’s buttercup
• Ranunculus muricatus – spinyfruit buttercup
• Ranunculus occidentalis – western buttercup
• Ranunculus orthorhynchus – straightbeak buttercup
• Ranunculus papulentus – large river buttercup
• Ranunculus parviflorus – smallflower buttercup
• Ranunculus pedatifidus – birdfoot buttercup
• Ranunculus peltatus – pond water crowfoot
• Ranunculus pensylvanicus – Pennsylvania buttercup
• Ranunculus platanifolius – large white buttercup
• Ranunculus populago – popular buttercup
• Ranunculus pusillus – low spearwort
• Ranunculus pygmaeus – pygmy buttercup
• Ranunculus recurvatus – hooked crowfoot
• Ranunculus repens – creeping buttercup
• Ranunculus rionii – water crowfoot
• Ranunculus sardous – hairy buttercup, Sardinian buttercup
• Ranunculus sceleratus – celery-leaved buttercup
• Ranunculus septentrionalis – swamp buttercup
• Ranunculus sieboldii
• Ranunculus testiculatus – bur buttercup
• Ranunculus thora – Thora buttercup
• Ranunculus trichophyllus Chaix ex Vill. – Thora buttercup
• Ranunculus uncinatus – woodland buttercup

Gardenia

Gardenias are waxy, white and very fragrant flowers Gardenias are one of the most popular exotic flowers

Gardenias are native to the tropical and subtropical regions of Africa, southern Asia, Australasia and Oceania. This species can be difficult to grow elsewhere because it demands high humidity to thrive and bright (not direct) light. Some types of gardenias can be grown as houseplants.

Kingdom – Plantae

Division -Magnoliophyta

Class -Magnoliopsida

Order  -Gentianales

Family -Rubiaceae

Genus -Gardenia

Gardenias symbolize purity and sweetness. They indicate secret love. They convey joy. They tell the receiver “you are lovely”.

ome Interesting Facts About Gardenias

• Gardenias are grown for their beautiful foliage and they make great cut flowers.
• Gardenias flowers from about mid-spring to mid-summer, i.e., May through July
• Many of the gardenia species are strongly scented.
• The genus Gardenia is named after Alexander Garden, a physician in Charleston of South Carolina.
• Gardenias can be used as screens, hedges, borders or ground covers.
• In France, Gardenia is the traditional flower which men wear as boutonnieres.
• The most popular cultivated species is the Cape jasmine, native to China.
• Each gardenia flower is followed by 6-sided berries of rich orange-red with long, elegant stems.
• Gardenia thunbergia produces a woody fruit which has hard, angular seeds inside.

About Gardenia Flower and Plant

Gardenias are very fragrant creamy-white flowers with glossy, dark-green leaves. Gardenia flowers are solitary or in small clusters, white or pale yellow. The gardenia flowers are with a tubular-based corolla with 5-12 lobes petals from 5-12 cm diameter.

Gardenia plants are evergreen shrubs and small trees growing to 1-15 m tall. The Gardenia plant leaves are opposite or in whorls of three or four. They are dark green and glossy 5-50 cm long and 3-25 cm broad, with a leathery texture. They are simple, entire, hairless, with wavy margin.

 

Growing Gardenias

• Gardenia plants need high humidity.
• A loose, well-drained organic soil is recommended.
• For best results plant gardenias in full sun, partial shade, or shifting shade.
• Dig a hole twice the size of the root ball.
• Thoroughly mix one part soil and one part planting mix.
• Check that the top of the original root ball is slightly above the level of the surrounding soil.
• Take the soil mixture and make a gentle mound of soil sloping away from the plant so that the water drains away from the trunk.
• Remove any air pockets, if present.
• Water the plant with a root stimulator.

Gardenia Care

• Do not over-water gardenias.
• Fertilize with an acid fertilizer.
• Check for aphids, mealybugs, spider mites, thrips and scales.
• Also check gardenia plants for bud drop, a common ailment.

Lily – Lilium

 

 Lilium (members of which are true lilies) is a genus of herbaceous flowering plants growing from bulbs, all with large prominent flowers. Lilies are a group of flowering plants which are important in culture and literature in much of the world. Most species are native to the temperate northern hemisphere, though their range extends into the northern subtropics. Many other plants have “lily” in their common name but are not related to true lilie

 

Scientific classification
Kingdom: Plantae
(unranked): Angiosperms
(unranked): Monocots
Order: Liliales
Family: Liliaceae
Subfamily: Lilioideae
Genus: Lilium

Names

The botanic name Lilium is the Latin form and is a Linnaean name. The Latin name is derived from the Greek λείριον, leírion, generally assumed to refer to true, white lilies as exemplified by the Madonna lily.^[1] The word was borrowed from Coptic (dial. Fayyumic) hleri, from standard hreri, from Demotic hrry, from Egyptian hrṛt “flower”. Meillet maintains that both the Egyptian and the Greek word are possible loans from an extinct, substratum language of the Eastern Mediterranean. The Greeks also used the word κρῖνον, krīnon, albeit for non-white lilies.The term “lily” has in the past been applied to numerous flowering plants, often with only superficial resemblance to the true lily, including lotus, ranunculus, tulip, iris, anemone, agapanthus, zantedeschia, daylily, and others. All English translations of the Bible render the Hebrew shūshan, shōshan, shōshannā as “lily”, but the “lily among the thorns” of Song of Solomon, for instance, may be the honeysuckle.

Range

The range of lilies in the Old World extends across much of Europe, across most of Asia to Japan, south to India, and east to Indochina and the Philippines. In the New World they extend from southern Canada through much of the United States. They are commonly adapted to either woodland habitats, often montane, or sometimes to grassland habitats. A few can survive in marshland and epiphytes are known in tropical southeast Asia. In general they prefer moderately acidic or lime-free soils.Lilies are tall perennials ranging in height from 2–6 ft (60–180 cm). They form naked or tunicless scaly underground bulbs which are their overwintering organs. In some North American species the base of the bulb develops into rhizomes, on which numerous small bulbs are found. Some species develop stolons. Most bulbs are deeply buried, but a few species form bulbs near the soil surface. Many species form stem-roots. With these, the bulb grows naturally at some depth in the soil, and each year the new stem puts out adventitious roots above the bulb as it emerges from the soil. These roots are in addition to the basal roots that develop at the base of the bulb.The flowers are large, often fragrant, and come in a range of colours including whites, yellows, oranges, pinks, reds and purples. Markings include spots and brush strokes. The plants are late spring- or summer-flowering. Flowers are borne in racemes or umbels at the tip of the stem, with six tepals spreading or reflexed, to give flowers varying from funnel shape to a “Turk’s cap”. The tepals are free from each other, and bear a nectary at the base of each flower. The ovary is ‘superior’, borne above the point of attachment of the anthers. The fruit is a three-celled capsule.Seeds ripen in late summer. They exhibit varying and sometimes complex germination patterns, many adapted to cool temperate climates.Naturally most cool temperate species are deciduous and dormant in winter in their native environment. But a few species which distribute in hot summer and mild winter area (Lilium candidum, Lilium catesbaei,Lilium longiflorum) lose leaves and remain relatively short dormant in Summer or Autumn, sprout from Autumn to winter, forming dwarf stem bearing a basal rosette of leaves until accept enough chilling requirement, the stem begins to elongate while warming.

Ecology

Lilies are used as food plants by the larvae of some Lepidoptera species including the Dun-bar.

Toxicity

Some Lilium species are toxic to cats. This is known to be so especially for L. longiflorum though other Lilium and the unrelated Hemerocallis can also cause the same symptoms. The true mechanism of toxicity is undetermined, but it involves damage to the renal tubular epithelium (composing the substance of the kidney and secreting, collecting, and conducting urine), which can cause acute renal failure. Veterinary help should be sought, as a matter of urgency, for any cat that is suspected of eating any part of a lily – including licking pollen that may have brushed onto its coat.

Cultivation

Many species are widely grown in the garden in temperate and sub-tropical regions. They may also be grown as potted plants. Numerous ornamental hybrids have been developed. They can be used in herbaceous borders, woodland and shrub plantings, and as patio plants. Some lilies, especially Lilium longiflorum, form important cut flower crops. These may be forced for particular markets; for instance, L. longiflorum for the Easter trade, when it may be called the Easter lily.Lilies are usually planted as bulbs in the dormant season. They are best planted in a south-facing, slightly sloping aspect, in sun or part shade, at a depth 2½ times the height of the bulb (except L. candidum which should be planted at the surface). Most prefer a porous, loamy soil, and good drainage is essential. Most species bloom in July or August. The flowering periods of certain lily species begin in late spring, while others bloom in late summer or early autumn.^[8] They have contractile roots which pull the plant down to the correct depth, therefore it is better to plant them too shallowly than too deep. A soil pH of around 6.5 is generally safe. The soil should be well-drained, and plants must be kept watered during the growing season. Some plants have strong wiry stems, but those with heavy flower heads may need staking.

Stargazer

Numerous forms, mostly hybrids, are grown for the garden. They vary according to the species and interspecific hybrids that they derived from, and are classified in the following broad groups:

• Asiatic hybrids (Division I). These are plants with medium sized, upright or outward facing flowers, mostly unscented. They are derived from central and East Asian species and interspecific hybrids, including L. amabile, L. bulbiferum, L. callosum, L. cernuum, L. concolor, L. dauricum, L. davidii, L. × hollandicum, L. lancifolium (syn. L. tigrinum), L. lankongense, L. leichtlinii, L. × maculatum, L. pumilum, L. × scottiae, L. wardii and L. wilsonii.
• Martagon hybrids (Division II). These are based on L. × dalhansonii, L. hansonii, L. martagon, L. medeoloides and L. tsingtauense. The flowers are nodding, Turk’s cap style (with the petals strongly recurved).
• Candidum (Euro-Caucasian) hybrids (Division III). This includes mostly European species: L. candidum, L. chalcedonicum, L. kesselringianum, L. monadelphum, L. pomponium, L. pyrenaicum and L. × testaceum.
• American hybrids (Division IV). These are mostly taller growing forms, originally derived from L. bolanderi, L. × burbankii, L. canadense, L. columbianum, L. grayi, L. humboldtii, L. kelleyanum, L. kelloggii, L. maritimum, L. michauxii, L. michiganense, L. occidentale, L. × pardaboldtii, L. pardalinum, L. parryi, L. parvum, L. philadelphicum, L. pitkinense, L. superbum, L. ollmeri, L. washingtonianum and L. wigginsii. Many are clump-forming perennials with rhizomatous rootstocks.
• Longiflorum hybrids (Division V). These are cultivated forms of this species and its subspecies. They are most important as plants for cut flowers, and are less often grown in the garden than other hybrids.
• Trumpet lilies (Division VI), including Aurelian hybrids (with L. henryi). This group includes hybrids of many Asiatic species and their interspecific hybrids, including L. × aurelianense, L. brownii, L. × centigale, L. henryi, L. × imperiale, L. × kewense, L. leucanthum, L. regale, L. rosthornii, L. sargentiae, L. sulphureum and L. × sulphurgale. The flowers are trumpet shaped, facing outward or somewhat downward, and tend to be strongly fragrant, often especially night-fragrant.
• Oriental hybrids (Division VII). These are based on hybrids L. auratum and L. speciosum, together with crossbreeds from several species native to Japan, including L. nobilissimum, L. rubellum, L. alexandrae, and L. japonicum. They are fragrant, and the flowers tend to be outward facing. Plants tend to be tall, and the flowers may be quite large. The whole group are sometimes referred to as “stargazers” because they appear to look upwards. (For the specific cultivar, see Lilium ‘Stargazer’.)
• Other hybrids (Division VIII). Includes all other garden hybrids.
• Species (Division IX). All natural species and naturally occurring forms are included in this group.

And there are subdivisions classified by flower aspect and form. EX:(flower aspect/flower form)Flower aspect:

• a up-facing
• b out-facing
• c down-facing

Flower form:

• a trumpet-shaped
• b bowl-shaped
• c flat (or with tepal tips recurved)
• d tepals strongly recurved (with the Turk’s cap form as the ultimate state)

Many newer commercial varieties are developed by using new technologies such as in vitro pollination, ovary culture and embryo rescue.

Pests and diseases

Aphids may infest plants. Leatherjackets feed on the roots. Larvae of the Scarlet lily beetle can cause serious damage to the stems and leaves. The scarlet beetle lays its eggs and completes its life cycle only on true lilies. Oriental, rubrum, tiger and trumpet lilies as well as Oriental trumpets (orienpets) and Turk’s cap lilies and native North American lily species are all vulnerable, but the beetle prefers some types. The beetle could also be having an effect on native Canada species and some rare and endangered species found in northeastern North America. Day lilies are excluded from this category. Plants can suffer from damage caused by mice, deer and squirrels. Slugs, snails and millipedes attack seedlings, leaves and flowers. Brown spots on damp leaves may signal botrytis (also known as lily disease). Various fungal and viral diseases can cause mottling of leaves and stunting of growth.

Propagation and growth

Lilies can be propagated in several ways;

• by division of the bulbs
• by growing-on bulbils which are adventitious bulbs formed on the stem
• by scaling, for which whole scales are detached from the bulb and planted to form a new bulb
• by seed; there are many seed germination patterns, which can be complex
• by micropropagation techniques (which include tissue culture);commercial quantities of lilies are often propagated in vitro and then planted out to grow into plants large enough to sell.

According to a study done by Anna Pobudkiewicz and Jadwiga the use of flurprimidol foliar spray helps aid in the limitation of stem elongation in oriental lilies.

Culinary and herb uses

China

Lilium bulbs are starchy and edible as root vegetables, although bulbs of some species may be very bitter. The non-bitter bulbs of L. lancifolium, L. pumilum, and especially L. brownii (Chinese: 百合 干; pinyin: bǎihé gān) and Lilium davidii var unicolor are grown on a large scale in China as a luxury or health food, and are most often sold in dry form for herb, the fresh form often appears with other vegetables. The dried bulbs are commonly used in the south to flavor soup. Lily flowers are also said to be efficacious in pulmonary affections, and to have tonic properties. Lily flowers and bulbs are eaten especially in the summer, for their perceived ability to reduce internal heat. They may be reconstituted and stir-fried, grated and used to thicken soup, or processed to extract starch. Their texture and taste draw comparisons with the potato, although the individual bulb scales are much smaller. There are also species which are meant to be suitable for culinary and/or herb uses. There are five traditional lily species whose bulbs are certified and classified as “vegetable and non-staple foodstuffs” on the National geographical indication product list of China.

Culinary use:

野百合Lilium brownii, 百合（变种）Lilium brownii var. viridulum, 渥丹 Lilium concolor, 毛百合 Lilium dauricum, 川百合 Lilium davidii, 东北百合 Lilium distichum, 卷丹 Lilium lancifolium, 新疆百合（变种） Lilium martagon var. pilosiusculum, 山丹 Lilium pumilum, 南川百合 Lilium rosthornii, 药百合（变种） Lilium speciosum var. gloriosoides.

Herb use:

野百合 Lilium brownii, 百合（变种）Lilium brownii var. viridulum, 渥丹 Lilium concolor, 毛百合 Lilium dauricum, 卷丹 Lilium lancifolium, 山丹 Lilium pumilum, 南川百合 Lilium rosthornii, 药百合（变种） Lilium speciosum var. gloriosoides, 淡黄花百合 Lilium sulphureum.

And there are researches about the selection of new varieties of edible lilies from the horticultural cultivars, such as ‘Batistero’ and ‘California’ among 15 lilies in Beijing, and ‘Prato’ and ‘Small foreigners’ among 13 lilies in Ningbo.

Japan

Yuri-ne (lily-root) is also common in Japanese cuisine, especially as an ingredient of chawan-mushi (savoury egg custard). The major lilium species cultivated as vegetable are L. leichtlinii var. maximowiczii, L. lancifolium, and L. auratum

Taiwan

The parts of lilium species which are officially listed as food material are the flower and bulbs of Lilium lancifolium Thunb., Lilium brownii F. E. Brown var. viridulum Baker, Lilium pumilum DC., Lilium candidum Loureiro. Most edible lily bulbs which can be purchased in a market are mostly imported from mainland China (only in the scale form, and most marked as 蘭州百合 Lilium davidii var unicolor) and Japan (whole bulbs, should mostly be L. leichtlinii var. maximowiczii). There are already commercially available organic growing and normal growing edible lily bulbs. The varieties are selected by the Taiwanese Department of Agriculture from the Asiatic lily cultivars that are imported from the Netherlands; the seedling bulbs must be imported from the Netherlands every year.

South Korea

The lilium species which are officially listed as herbs are 이 약은 참나리 Lilium lancifolium Thunberg; 백합 Lilium brownii var. viridulun Baker; 또는 큰솔나리 Lilium pumilum DC.

The “lily” flower buds known as jīnzhēn (金针, “golden needles”) in Chinese cuisine are actually from the daylily Hemerocallis fulva.

Taxonomy

Taxonomical division in sections follows the classical division of Comber, species acceptance follows the World Checklist of Selected Plant Families, the taxonomy of section Pseudolirium is from the Flora of North America, the taxonomy of Section Liriotypus is given in consideration of Resetnik et al. 2007, the taxonomy of Chinese species (various sections) follows the Flora of China  and the taxonomy of Section Archelirion follows Nishikawa et al. as does the taxonomy of Section Archelirion.

There are seven sections;

• Martagon
• Pseudolirium
• Liriotypus
• Archelirion
• Sinomartagon
• Leucolirion
• Daurolirion

Some species formerly included within this genus have now been placed in other genera. These genera include Cardiocrinum, Notholirion, Nomocharis and Fritillaria.